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Lee ATK, Macray MB, Ryan PG, Alexander GJ 2021. Tortoise mortality along fence lines in the Karoo region of South Africa. Journal for Nature Conservation 59
Alström P, Barnes KN, Olsson U, Barker FK, Bloomer P, Khang AA, Qureshi MA, Guillaumet A, Crochet P-A, Ryan PG 2013. Multilocus phylogeny of the avian family Alaudidae (larks) reveals complex morphological evolution, non-monophyletic genera and hidden species diversity. Molecular Phylogenetics and Evolution 69 (3) 1043-1056
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