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Lee ATK, Macray MB, Ryan PG, Alexander GJ 2021. Tortoise mortality along fence lines in the Karoo region of South Africa. Journal for Nature Conservation 59

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Sherley RB, Crawford RJM, Dyer BM, Kemper J, Makhado AB, Masotla M, Pichegru L, Pistorius PA, Roux J-P, Ryan PG, Tom D, Upfold L, Winker H 2019. The status and conservation of the Cape Gannet Morus capensis. Ostrich 90 (4) 335-346

Crawford RJM, Ryan PG, Williams AJ 1991. Seabird consumption and production in the Benguela and Western Agulhas ecosystems. South African Journal of Marine Science 11 357-375

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