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Crawford RJM, Makhado AB, Whittington PA, Randall RM, Oosthuizen WH, Waller LJ 2015. A changing distribution of seabirds in South Africa—the possible impact of climate and its consequences. Frontiers in Ecology and the Environment Review Article

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Whittington PA, Randall RM, Randall BM, Wolfaardt AC, Crawford RJM, Klages NTW, Bartlett PA, Chesselet YJ, Jones R 2005. Patterns of movements of the African penguin in South Africa and Namibia. African Journal of Marine Science 27 (1) 215-229

Whittington PA, Randall RM, Crawford RJM, Wolfaardt AC, Klages NTW, Randall BM, Bartlett PA, Chesselet YJ, Jones R 2005. Patterns of immigration to and emigration from breeding colonies by African penguins. African Journal of Marine Science 27 (1) 205-213

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