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Sherley RB, Makhado AB, Crawford RJM, Hagen C, Kemper J, Ludynia K, Masotla MJ, McInnes A, Pichegru L, Tom D, Upfold L, Waller LJ 2024. The African Penguin Spheniscus demersus should be considered Critically Endangered. Ostrich 95 (3)
Vanstreels RET, Parsons NJ, McGeorge C, Hurtado R, Ludynia K, Waller L, Ruthenberg M, Purves A, Pichegru L, Pistorius PA 2019. Identification of land predators of African Penguins Spheniscus demersus through post-mortem examination. Ostrich 90 (4) 359-372
Kohler SA, Connan M, Hill JM, Mablouké C, Bonnevie B, Ludynia K, Kemper J, Huisanem J, Underhill LG, Cherel Y, McQuiad CD, Jaquemet S 2011. Geographic variation in the trophic ecology of an avian rocky shore predator, the African black oystercatcher, along the southern African coastline. Marine Ecology Progress Series 435 235-249

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