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Ras V, Neethling S, Engelbrecht A, Morandini AC, Bayha KM, Skrypzeck H, Gibbons MJ 2020. There are three species of Chrysaora (Scyphozoa: Discomedusae) in the Benguela upwelling ecosystem, not two. Zootaxa 4778 (3) 401-438
Flynn BA, Richardson AJ, Brierley AS, Boyer DC, Axelsen BE, Scott L, Moroff NE, Kainge PI, Tjizoo BM, Gibbons MJ 2012. Temporal and spatial patterns in the abundance of jellyfish in the northern Benguela upwelling ecosystem and their link to thwarted pelagic fishery recovery.   African Journal of Marine Science 34 (1) 131-146

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