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Frolov AV, Akhmetova LA, Scholtz CH 2008. Revision of the obligate mushroom‐feeding African "dung beetle" genus Coptorhina Hope (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of Natural History 42 (21-22) 1477-1508

Website link:http://www.tandfonline.com/doi/full/10.1080/00222930802002535#.VLYnUGctApE

Frolov AV, Scholtz CH 2003. A new genus and species of dung beetle from southern Namibia (Coleoptera: Scarabaeidae: Scarabaeinae). African Entomology 11 (2) 297-299

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A new genus and species of dung beetle from southern Namibia_2003.pdf	800.7 KB

Holm E, Scholtz CH 1980. Structure and pattern of the Namib Desert dune ecosystem at Gobabeb. Madoqua 12 (1) 5-39

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Structure and patterns of the Namib Desert dune ecosystem_Gobabeb.pdf	3.66 MB

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