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Lockley MG, Helm CW, Cawthra HC, de Vynck JC, Perrin MR 2021. Pleistocene golden mole and sand-swimming trace fossils from the Cape coast of South Africa. Quaternary Research 101 169-186
Perrin MR, Boyer DC 2000. Seasonal changes in the population dynamics of hairy-footed gerbils in the Namib Desert. South African Journal of Wildlife Research 30 (2) 73-84
Perrin MR, Dempster ER, Downs CT, Boyer DC 1999. Gerbillurus tytonis. Mammalian Species 607 1-4
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Perrin MR, Boyer DC 1996. Habitat selection by the gerbils Gerbillurus paeba and G. tytonis in the Namib Desert dunes. South African Journal of Wildlife Research 26 (3) 71-76

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