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Bauer H, Packer C, Funston PF, Henschel P, Nowell K 2016. Panthera leo. The IUCN Red List of Threatened Species 2016: e.T15951A115130419.
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Miller JRN, Balme G, Lindsey PA, Loveridge AJ, Becker MS, Begg C, Brink H, Dolrenry S, Hunt JE, Jansson I, MacDonald DW, Mandisodza-Chikerema RL, Cotterill AO, Packer C, Rosengren D, Stratford K, Trinkel M, White PA, Winterbach C, Winterbach HEK, Funston PJ 2016. Aging traits and sustainable trophy hunting of African lions. Biological Conservation (201) 160-168
Barthold JA, Loveridge AJ, MacDonald DW, Packer C, Colchero F 2016. Bayesian estimates of male and female African lion mortality for future use in population management. Journal of Applied Ecology (53) 295–304
Packer C, Scheel D, Pusey AE 1990. Why lions form groups: Food is not enough. The American Naturalist 136 (1) 1-19

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