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Uphyrkina O, Johnson WE, Quigley H, Miquelle D, Marker L, Bush M, O'Brien SJ 2001. Phylogenetics, genome diversity and origin of modern leopard, Panthera pardus. Molecular Ecology 10 2617–2633
Miththapala S, Seidensticker J, O'Brien SJ 1996. Phylogeographical subspecies recognition in leopards (Panthera pardus): molecular genetic variation. Conservation Biology 10 (4) 1115-1131
Brown EW, Miththapala S, O'Brien SJ 1993. Prevalence of exposure to feline Immunodeficiency Virus in exotic felid species. Journal of Zoo and Wildlife Medicine 24 (3) 357-364

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