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Turner WC, Küsters M, Versfeld W, Horak IG 2017. Ixodid tick diversity on wild mammals, birds, and reptiles in and around Etosha National Park, Namibia. African Journal of Ecology 55 (4) 714-721
Penzhorn BL, Oosthuizen MC, Bosman A-M, Kilian JW, Horak IG 2008. Black Rhinoceros (Diceros bicornis) populations in northwestern Namibia are apparently not infected with Piroplasms. Journal of Wildlife Diseases 44 (4) 1032-1035
Knapp SE, Krecek RC, Horak IG, Penzhorn BL 1997. Helminths and arthropods of black and white rhinoceroses in southern Africa. Journal of Wildlife Diseases 33 (3) 492-205

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