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van Jaarsveld AS, Henschel JR, Skinner JD 1987. Improved age estimation in spotted hyaenas (Crocuta crocuta). Journal of Zoology 213 (4) 758-762
Tilson RL, Henschel JR 1986. Spatial arrangement of spotted hyaena groups in a desert environment, Namibia. African Journal of Ecology 24 (3) 173-180

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