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Mucina L, Jürgens N, Le Roux A, Rutherford MC, Schmiedel U, Esler KJ, Powrie LW, Desmet PG, Milton SJ 2006. Succulent Karoo Biome. Strelitzia 19 221-299
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Midgley JJ, Cowling RM, Hendricks H, Desmet PG, Esler K, Rundel P 1997. Population ecology of tree succulents (Aloe and Pachypodium) in the arid western Cape: decline of keystone species. Biodiversity and Conservation 6 (6) 869-876

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