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Cowling RM, Esler KJ, Rundel PW 1999. Namaqualand, South Africa – an overview of a unique winter-rainfall desert ecosystem. Plant Ecology 142 (1-2) 3-21
Hoffman MT, Cowling RM, Douie C, Pierce SM 1989. Seed predation and germination of Acacia erioloba in the Kuiseb River Valley, Namib Desert. South African Journal of Botany 55 (1) 103-106

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