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Cowling RM, Esler KJ, Rundel PW 1999. Namaqualand, South Africa – an overview of a unique winter-rainfall desert ecosystem. Plant Ecology 142 (1-2) 3-21
Cowling RM, Hilton-Taylor C 1991. 4 - Plant biogeography, endemism and diversity. The karoo - Ecological Patterns and Processes 42-46

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