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Desmet PG, Cowling RM, Ellis AG, Pressey RL 2002. Integrating biosystematic data into conservation planning: Perspectives from southern Africa's Succulent Karoo. Systematic Biology 51 (2) 317–330
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Cowling RM, Dlamini TS, Stock WD 1999. Plant Induced Fertile Islands as Possible Indicators of Desertification in a Succulent Desert Ecosystem in Northern Namaqualand, South Africa. Plant Ecology 142 (1/2 The Plant Ecology of Namaqualand, South Africa (Jun., 1999)) 161-167
Midgley JJ, Cowling RM, Hendricks H, Desmet PG, Esler K, Rundel P 1997. Population ecology of tree succulents (Aloe and Pachypodium) in the arid western Cape: decline of keystone species. Biodiversity and Conservation 6 (6) 869-876

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