Basal lineages in these two taxa are present in the southwestern Cape of South Africa where several species of Pachydactylus also occur, but their chief diversification has been in the temperate Cape Fold Mountains (Goggia) and along the Eastern Escarpment and into the Eastern Cape (Afroedura), where Pachydactylus is poorly represented. Conversely, Pachydactylus and its related genera have their greatest diversity in the arid regions of the Northern Cape and Namibia, where the other genera are represented by only two species each. The approximately contemporaneous diversification of these genera suggests that ecological interactions between them may have contributed to their largely complementary distributions.