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Hattingh J, Ganhao M, Naidu SG 1984. Haemolymph composition in Namib Desert tenebrionid beetles. Comparative Biochemistry and Physiology Part A: Physiology 78 (3) 541-545
Naidu SG 2001. Osmoregulation in Onymacris rugatipennis, a free-ranging tenebrionid beetle from the Namib Desert. Comparative Biochemistry and Physiology Part A: Molecular & Integrative Physiology 129 (4) 873-885

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