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Lewin A, Feldman A, Bauer AM, Belmaker J, Broadley DG, Chirio L, Itescu Y, LeBreton M, Maza E, Meirte D, Nagy ZT, Novosolov M, Roll U, Tallowin O, Trape J, Vidan E, Meiri S 2016. Patterns of species richness, endemism and environmental gradients of African reptiles. Journal of Biogeography 43 (12) 2380-2390
Branch WR, Baard EHW, Haacke WD, Jacobsen N, Poynton JC, Broadley DG 1988. A provisional and annotated checklist of the Herpetofauna of southern Africa. The Journal of the Herpetological Association of Africa 34 (1) 1-19
Poynton JC, Broadley DG 1988. Amphibia Zambesiaca 4. Bufonidae. Annals of the Natal Museum (29) 447-490
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